Richard Arthur Studhalter

1887 – 1958

• Germination of spores and development of juvenile thallus of Riella americana
• Independence of sporophyte in Riella and Sphaerocarpus
• The foot of Riella americana and its relation to nutrition of the sporophyte
• The gemmaling of Riella Americana
• The lateral leaf scale of Riella americana
• The ventral scale of Riella americana